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I have often seen Hamster's Bruxing in sleeping. But my hamster Comy does it every time when I get a massage.
I was curious what her actions meant, so I searched Google.
And I learned about rats Bruxing.
Rats do that when they feel comfortable.
I think, it is also like a cat growling. Cats growl when they feel good. But even when Cat feel sick and unpleasant, Cat growl.
Even hamsters seem to have Bruxing when they feel bad.