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Despite their rather minor status in modern times, the sloths were once a highly diverse and successful group of mammals. A part of the endemic South American order Xenarthra, sloths first emerge in the fossil record during the Early Oligocene but almost certainly diverged before this. Developing into browsing and grazing herbivores, an early lineage of sloths, the Megalocnids, successfully colonized the Caribbean and survived there until roughly 5000 years ago. All more derived sloths belong to two major clades: the Mylodontoids and the Megatherioids. Both of these lineages greatly diversified, with genera entering North America during the Late Miocene before the Great American Interchange. Some forms such as Megatherium and Eremotherium developed massive body sizes, weighing as much as modern elephants, while others took to the seas to exploit a boom in sea grasses. Despite possessing a range that covered the entirety of both American continents by the Pleistocene, the mainland ground sloths all died out by the Early Holocene c. 10,000 years ago. This was due to a mixture of climate change and human hunting, leave only two small arboreal genera to persist into modern times.
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