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In the year 2387, the galaxy is threatened by an unusually volatile supernova. Ambassador Spock pilots a ship carrying "red matter", which will create an artificial black hole to consume the supernova. Before Spock completes his mission, the supernova destroys the planet Romulus. Captain Nero of the Romulan mining ship Narada attempts to exact revenge on Spock, whom he blames for the destruction of his homeworld and its inhabitants, including his wife and unborn child. Both ships, however, are pulled in to the black hole's event horizon and travel into the past.
The Narada arrives first, 154 years in the past. Its arrival is preceded by what looks like a lightning storm in space, which the Federation starship USS Kelvin investigates. Nero attacks the Kelvin, causing severe damage. Acting-Captain George Kirk orders an evacuation and stays behind to provide cover for the fleeing shuttlecraft, his pregnant wife among the passengers. George pilots the Kelvin into the Narada moments after his son James Tiberius Kirk is born. Spock arrives 25 years later, where he is captured by Nero and marooned on the planet Delta Vega so that he can witness the destruction of Vulcan, as retribution for Spock's failure to save Romulus.
Kirk grows up to be an intelligent, though reckless and cynical, young man. A young Spock grows up on Vulcan and is discriminated against because of his half-human heritage. To get Kirk to embrace his potential, Captain Christopher Pike challenges Kirk to emulate his father's heroism and convinces him to join Starfleet. En route to Starfleet Academy, Kirk befriends fellow cadet Leonard McCoy. In his third year at the Academy, Kirk is accused of cheating on the Kobayashi Maru test by its programmer, Commander Spock. At the resultant hearing, Starfleet receives a distress signal from Vulcan that indicates the appearance of a lightning storm in space, and with the primary fleet engaged in an operation, the cadets are mobilized to help crew the ships in orbit. McCoy smuggles the grounded Kirk aboard the USS Enterprise as a patient under his care.
Kirk recognizes the similarities between the Vulcan incident and the encounter that destroyed the Kelvin, and warns Pike that the fleet is heading into a trap. The Enterprise arrives late to find the fleet destroyed and the Narada drilling into Vulcan's core. Nero orders Pike to surrender himself via shuttlecraft. Pike obeys, promoting Spock to captain and Kirk to first officer. En route to the Narada, Kirk, Hikaru Sulu, and Chief Engineer Olson perform an orbital skydive onto the drilling platform to destroy it; Olson is killed, but Kirk and Sulu stop the drill. However, it has drilled deep enough for Nero to launch red matter into the planet's core, imploding the planet into a black hole. Spock rescues some of the planet's elders, including his father, but his mother dies along with the majority of the planet's population. Nero sets course for Earth and tortures Pike for the command codes to its perimeter defenses.
After a heated argument with Spock about their next move, Kirk is marooned on Delta Vega for mutiny. Spock orders the ship to rendezvous with the rest of the fleet. On Delta Vega, Kirk encounters Ambassador Spock, who relays the future's events through a mind meld and insists that Kirk must become captain of the Enterprise. The two travel to a nearby Starfleet outpost where they meet Montgomery Scott. The elder Spock beams Kirk and Scott aboard the Enterprise. Once aboard, Kirk deliberately enrages Spock to force him to acknowledge that he is emotionally compromised, thereby forfeiting command which then passes to Kirk.